Referred specimens:
(Text-figs. 14E, J, 15D).
Subfamily Pterodonti~ae PoLLY, 1996
Genus Hyainailouros BiiDERMANN, 1863
Hyainailouros, species tdeteterminae
. (Text-figs.l4, 15)
KNM-LS 18215, """""'' of riglrt low< ! ,,ru.,, (Ten-fig. 15B}; KNM -LS 18230, rii;h< m3 pm<ooo.Ud
Lo c alit i e s : All pterodontine material is from Losodok. . .,
Measurements: The most impressive feature of material refera~le to Hyainailouros or Megistotherium in Africa is simply its
gigantic size. The Losodok m3 has a crown height as measured on the bu cal side of 31 mm, and 32.3 mm as measured to the base on the
lingual side. The width of the crown at its base is >19 mm. The length of he preserved fragment (minus the paraconid) is 32 mm. This is
significantly smaller than the earliest Miocene specimen of a Hyainailouros like creodont in East Africa (a Meswa Bridge specimen; KNMME
22);. that specimen has a crown height of 40.7 mm and a crown widt of > 26 mm. The longest mesiodistal diameter of the Losodok
canine fragment is 32.8 mm, and the buccolingual diameter is 21.9 mm. T e comparable measurements of the MeswaBridge specimen are
40.9 and 27.2 mm. In contrast, the Losodok specimen is significantly larger than the undescribed tooth fragments of a large, Hyainailouroslike
creodont from Chilga, Ethiopia, the next oldest record of such creot!, nts in the African fossilrecord
D e s c rip ti on: The fragment of m3 represents a huge tooth. The mesial part of the tooth· is missing,
anterior to the notch separating paraconid from protoconid. The protoconid is simple and sectorial. It bears a
carnassial wear facet on the buccal side of its leading edge.l Lingual to this leading edge is a groove dropping
down to the lingual fossa of the paraconid-protoconid note ·. The distal blade of the protoconid is a sharp crest
without a wear facet. On the lingual side of this distal blade ·s afurrow running down the tooth to a distolingual
cingulum. Situated on the distal tooth border is a small s lid. The other specimen, the canine segment, has a
distinct, sharp keel on the upper (distal) surface. There is a l ngitudinal groove on the lingual face. The tooth is
smoothly rounded buccally and mesially. In these respects, i matches closely with the lower canines of an early
Miocene ~riant nterodontine from Meswa Brid~re (KNM-M 12 and 20). Discussioh: The newrmaterial c~ bealloca eel toHyainatlou~os,pending work on some of the better .
. prese~ed Miocene specimens that pelongtqthisgro p. An e¥1Y Miocenespedmenfrom Meswa Bridge should
help resolve the classification of giantpterodontines f omEast Africa: it resemblesNorth African Megistotherium
but differs from Hyainailouros in having . a disti ct, trenchant hypoconid on the p4, but it differs from
Megistotherium in having a two-rooted m1, and h inference; it presumably retained a typical m1 crown; in
contrast the mlcrownof Megistotherium is asimpl rounded knob vrith a single root (RAsMUSSEN et al. 1989).
The size increase from Chilga (26-27 Ma) to Los dok (24..:.26 Ma) to Meswa Bridge (19-22 Ma) suggests the
simple hypothesis that Hyainailouros was evolving 1 ger size over this time interval,\perhaps in response to the
increasing size of prohoscidean or anthracothere pre . . . . . .
In addition to the material described above, there are two other Losodok specimens of probable pterodontirie
creodonts. One of these is a right lower premolar in a jaw fragment that is similar to the· premolars of Pterodon
· (KNM-LS 18218); the other is an unidentifiable pre olar crown (KNM~LS 18213) .         Text-fig. 15. Dental specimens of Hyainailouros sp. from Losodok to Meswa Bridge specimens of Hyainailouros-type creodonts.
A, KNM-ME 12, rightlower canine from Meswa Bridge, next to B, KNM-LS 18215, fragment of a right lower canine from Losodok C,
KNM-ME 22, right lower m1 from Meswa Bridge, compared to D, KNM-ts 18230, right lower m3 from Losodok. The canine segment has
a distinct, sharp keel on the upper (distal) surface. The tooth is smoothly rofnded buccally and mesially. In these respects, it matches closely
with the lower canines of the early Miocene giant pterodontine from MesFa Bridge (KNM-ME 12). Entire two-toned scale bar= 1 em.

https://www.app.pan.pl/archive/publishe … 34-125.pdf  BRIGITTE LANGE-BADRE and DEMBERLYIN DASHZEVEG
ON SOME OLIGOCENE CARNIVOROUS MAMMALS
FROM CENTRAL ASIA   

черепа птеродонтин , азиатский эпиптеродон по форме скуловых дуг больше всех похож на череп мегистотерия.

JUL 2 6 1935
BULLETIN
OF THE
MUSEUM OF COMPARATIVE ZOOLOGY     https://84a69b9b8cf67b1fcf87220d0dabdda … printed%20(HD).pdf    Рома ,что скажешь по этим находкам гемипсалодона ,и он ли это ?

http://journals.plos.org/plosone/articl … ne.0135698  A New Large Hyainailourine from the Bartonian of Europe and Its Bearings on the Evolution and Ecology of Massive Hyaenodonts (Mammalia)   Классное название дали- церберус .

Мегистотерия уменьшили до 500 кг - https://en.wikipedia.org/wiki/Megistotherium

Ничего там еще не исправлено Сан Саныч.
500 кг - это, млин, голова на ножках какая-то получается

Velizar Simeonovski   
     Мне очень нравится эта картина ,помните находку диниктиса с пробитым черепом .предположительно гиенодоном хоридусом !  Хочется увидеть картину с мегистотерием

Machaeroides когда конкретно жил?

  Подогнал примитивно ,оба куска около 20 см длиной каждый выходит !

  Кто нибудь в курсе ,где есть описание покстрального скелета неогиенодона макроцефалус ? У Лаврова только описание черепа . Скелет находится в музее в Москве , у иностранцев с описаниями находок как то все проще - обидно !

  Кстати , гиенодон хоридус 1375 вполне мог быть самкой , у самцов более крупные черепа по отношению к всему телу !